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Nearshore Marine Fish Research Program

The programatic hierarchy



Research Scientists


Clare L. Wormald

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Research Scientist and Part-time Lecturer
Deparment of Biology
18111 Nordhoff Street
Northridge, CA 91330-8303
Office: 818-677-3340
Fax: 818-677-2034


2007 Ph.D. Natural Resources Science, University of Rhode Island, Rhode Island, USA.
1997 M.Sc. Tropical Coastal Management, University of Newcastle upon Tyne, UK.

Research Interests

I am interested in the processes that influence the distribution and the demographic rates of fish populations, including the influence of habitat, population density, and the effects of fishing and other anthropogenic impacts.

Photo: Kerry Nickols



White, J.W., Samhouri, J.F., Stier, A.C., Wormald, C.L., Hamilton, S.L. & Sandin, S.A. 2010. Synthesizing mechanisms of density dependence in reef fishes: behavior, habitat configuration, and observational scale. Ecology 91: 1949-1961.

Paddack, M.J., Reynolds, J.D., Aguilar, C, Appeldoorn, R.S., Beets, J., Burkett, E.W., Chittaro, P.M., Clarke, K., Esteves, R., Fonseca, A.C., Forrester, G.E., Friedlander, A.M., García-Sais, J., González-Sansón, G., Jordan, L.K.B., McClellan, D.B., Miller, M.W., Molloy, P.P., Mumby, P.J., Nagelkerken, I., Nemeth, M., Navas-Camacho, R., Pitt, J., Polunin, N.V.C., Reyes-Nivia, M.C., Robertson, D.R., Rodríguez-Ramírez, A., Salas, E., Smith, S.R., Spieler, R.E., Steele, M.A., Williams, I.D., Wormald, C.L., Watkinson, A.R. & Côté, I.M. 2009. Recent region-wide declines in Caribbean reef fish abundance. Current Biology 19: 590–595.

Wormald, C.L. & Steele, M.A. 2008. Testing assumptions of mark-re­capture theory in the coral reef fish Lutjanus apodus. Journal of Fish Biology 73: 498–509.

Wormald, C.L. 2007. Effects of density and habitat structure on growth and survival of harvested coral reef fishes. Ph.D. Dissertation, University of Rhode Island, Kingston, Rhode Island, USA.

Wormald, C.L. 1997. Recommendations for community based resource management at Danjugan Island, Negros Occidental, Philippines. M.Sc. Thesis, University of Newcastle-upon-Tyne, Newcastle-upon-Tyne, United Kingdom.

Prior, M., Ormond, R.F.G., Hitchen, R.G., & Wormald, C.L. 1995. The impact on natural resources of activity tourism: a case study of diving in Egypt. International Journal of Environmental Studies 48: 201-209.

Wormald, C.L. 1993. Behavioural assessment of divers and reef resource management strategies: Ras Mohammed National Park, Egypt. Honors Thesis, University of York, York, United Kingdom.